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Weisser (1975) stated that the type host may have been either Oryx dammah (Cretschmar) or Addax nasomaculatus (Blainville) (Artiodactyla: Bovidae). [ex. Durden, L. A., Musser, G. G. 1994. Bulletin of the American Museum of Natural History 218: 1-90]
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